ATHENS, Greece -- Five suspects implicated in the killing of a University of California, Berkeley professor appeared in court in Athens on Thursday, including his ex-wife, who has denied all charges.

Greek police have charged five people in the case of murdered UC Berkeley professor Przemyslaw Jeziorski, including his ex-wife and her current partner. Jeziorski was shot multiple times at close range on July 4 while visiting Athens to see his children and attend a family custody hearing.

Greek officials say the broad-daylight attack came one day after a custody hearing over the victim's 10-year-old twins.