ATHENS, Greece -- Five suspects implicated in the killing of a University of California, Berkeley professor appeared in court in Athens on Thursday, including his ex-wife, who has denied all charges.

The ex-wife of a University of California, Berkeley professor who was fatally shot during a visit to Greece earlier this month is among five people who have been charged in his killing, reports say.

Greek officials say the broad-daylight attack came one day after a custody hearing over the victim's 10-year-old twins.